Pterodactylus ("winged finger") is an extinct flying reptile genus of pterosaurs, whose members are popularly known as "pterodactyls". It is currently thought to contain only a single species, Pterodactylus antiquus, the first pterosaur species to be named and identified as a flying reptile.
It was a carnivore, and probably preyed upon fish and other small animals. Like all pterosaurs, Pterodactylus had wings formed by a skin and muscle membrane stretching from its elongated fourth finger to its hind limbs. It was supported internally by collagen fibres and externally by keratinous ridges.
Pterodactylus is known from over 30 fossil specimens, and though most of those are juveniles, many preserve complete skeletons. Pterodactylus antiquus was a relatively small pterosaur, with an estimated adult wingspan of about 1.04 meters (3 feet 5 inches) (the only known adult specimen is represented by an isolated skull). Other "species" were once thought to be smaller. However, these smaller specimens have been shown to represent juveniles of Pterodactylus, as well as its contemporary relatives including Ctenochasma, Germanodactylus, Aerodactylus, Aurorazhdarcho, and Gnathosaurus.
Pterodactylus, like related pterosaurs, had a crest on its skull composed mainly of soft tissues. In adult Pterodactylus, this crest extended between the back edge of the antorbital fenestra (the largest opening in the skull) and the back of the skull. In at least one specimen, the crest had a short bony base, also seen in related pterosaurs like Germanodactylus. Solid crests have only been found on large, fully adult specimens of Pterodactylus, indicating that this was a display structure that became larger and more well developed as individuals reached maturity. Bennett (2013) noted that other authors claimed that the soft tissue crest of Pterodactylus extended backward behind the skull; Bennett himself, however, didn't find any evidence for the crest extending past the back of the skull. Two specimens of P. antiquus (the holotype specimen BSP AS I 739 and the incomplete skull BMMS 7, the largest known skull of P. antiquus) have a low bony crest on their skulls; in BMMS 7 it is 47.5 mm long (more or less 24% of the estimated total length of its skull) and has a maximum height of 0.9 mm above the orbit.
The Land Before Time
Two distinct "breeds" of Pterodactylus appear in the series; with the first appearing in The Land Before Time, during Littlefoot's birth, wherein they are depicted with very Pteranodon-like head crests and skulls.
The second "breed" appears in The Land Before Time V: The Mysterious Island, in the form of a predatory "Sharptooth Flyer" that captures Ducky. They bear a greater resemblance to conventional Pterodactlyus than those in the original film.